Encyclopedia of Animal Behavior
- 3rd Edition - June 1, 2026
- Latest edition
- Editor: Benjamin Oldroyd
- Language: English
Encyclopedia of Animal Behavior, Third Edition, Four Volume Set is a unique curated collection of punchy essays on all aspects of animal behavior, written by experts and carefu… Read more
Encyclopedia of Animal Behavior, Third Edition, Four Volume Set is a unique curated collection of punchy essays on all aspects of animal behavior, written by experts and carefully edited for readability and accessibility. Animal behavior is usually defined as what animals do, i.e. their movements. We can ask how they move, (i.e. the physiological and neurological causes of movement) and why they move. To answer this latter question, behavioralists generally turn to evolutionary explanations. The Encyclopedia gives equal weight to these ‘how’ (proximate) and `why` (ultimate) explanations of animal behavior. The third edition updates and expands the previous version to cover all substantive topics in the field. Animal behavior is one of the largest subdisciplines in biology. Therefore, chapters focus on broad concepts and avoid becoming entrapped in taxon-specific detail. To illustrate this, one could consider the topic ‘cooperative hunting’. One way to cover this topic would be to have individual chapters on hunting in lions, mongoose, African wild dogs, army ants, dolphins etc. But it would serve the reader much better if a comparative approach was taken to see if there are commonalities across taxa. For example, is communication necessary for cooperative hunting? Is there any evidence of planning or is every hunt opportunistic? This comparative approach requires authors to provide a synthesis, pushing beyond specific taxa of expertise, but helps readers making useful connections. This is a fundamental feature of this work. This edition retains chapters on the historical development of the field, which is important to our understanding of where the discipline sits today, but similarly the work is forward leaning, including the authors` perspective of where their discipline is headed. Structured chapters and cross-references help readers following their interests with ease. Like a bird field guide, this encyclopedia is an entrée to deeper knowledge. Readers of the encyclopedia are encouraged to explore beyond individual chapters via links to related ones, key reviews and relevant online material.
- Comprehensive: covers the entire field of animal behavior, making the encyclopedia the go-to reference for getting up-to-speed on any substantive topic
- Conceptual: Each chapter focuses on broad concepts and avoids becoming entrapped in taxon-specific detail. This allows readers to make useful connections, beyond their direct specialty
- Authoritative: transparency over authorship and editorship means that the Encyclopaedia can be used by students and scientists as a reliable and citable reference
- Connected: structured chapters and cross-references allow readers to explore connected topics, and find pointers to further reading and online resources
Students and practising scientists in areas related to general biology, behavioral ecology, neuroscience, animal welfare, animal science, veterinary science, ethology
Cognition & Communication
Animal Arithmetic
Animal Tool use
Categories and Concepts: Language-Related Competences in Non-Linguistic Species
Chimpanzee and bonobo
Cognitive Development in Chimpanzees
Collaborative behavior
Conflict resolution
Deception: Competition by Misleading Behavior
Emotion and Social Cognition in Primates
Empathetic Behavior
Gestural communication in the great apes
Mental Time Travel: Can Animals Recall the Past and Plan for the Future?
Metacognition and Metamemory in Non-Human Animals
Non-Elemental Learning in Invertebrates
Primate Archaeology
Comparative Thanatology: Responses to the Dying and Dead, with Special Reference to Primates
Cold-Blooded Cognition: Cognition in Testudines
Collective decision making
Avian Tool Use in the Wild
Hemispherical asymetry in animal cogntion (across species)
Acoustical Signals and Signallers – in air and water.
Chemical signaling; air, water, and on the substrate
Electric signals
Multimodal signaling
Vibrational Signals: Sounds Transmitted Through Solids
Mating signals, including advertisement and courtship
Parent-offspring Signaling
Signals in conflict resolution: conventional signals, aggression and territoriality
Individual signatures in animal groups: cetaceans
Telling Your Friends Where the Goodies are – Recruitment Signals for Food and Habitat
Signals in Insect Social Organization
Communication networks
Adjustments to Facilitate Communication in Noisy Environments
Bird vocalizations and linguistics
Rhythm and music in animal signals
Elephant communication
The Evolution of Diverse Intelligence: Comparing Corvid and Cephalopod Cognition
Elephant cognition
Behavioral Ecology
Cooperative breeding
Ecology of Fear
Isolating Mechanisms and Speciation
Methodology: Cost and Benefit Analysis
Personality and Evolution
Quantitative genetics of behavior
Specialization
An Introduction to Optimal Foraging Theory
Habitat Selection
Hunger and Satiety: Linking Mechanisms Behavior, Ecology and Evolution
Patch Exploitation
Understanding movements of foraging animals – An approach based on Optimal Foraging Theory
Food hoarding
Internal Energy Storage
How variance and risk affect foraging behaviour
Aposematism as a defence against predation
Foraging in Groups
Plant-pollinator co-evolution & Optimal Foraging Theory
Ectoparasite Behavior
Parasite-Induced Behavioral Change: Mechanisms
Avoidance of Parasites
Animal Medication: The Passive Prevention and Active Treatment of self and others
Links between Nutrition, Immunity and Infection
Reproductive Behavior and Parasites: Vertebrates
Social Behavior and Parasites
Migration in insects
Predator Evasion
Predator Avoidance: Mechanisms
Defensive Avoidance
Defensive Coloration
Group living
Anti Predatory Vigilance
Risk Allocation in Anti Predator Behavior
Defensive Chemicals
Predation risk and life histories
Defensive risk taking in Animals
Games Played by Predators and Prey
Intraguild Predation
Antipredator Benefits from Heterospecifics
Parasitoids
Crustacean Social Evolution
Microbes: Social Evolution
The Honey Bee's Dance Language
Collective Intelligence in Social Animals
Nest Site Choice in Social Insects
Group Movement
Parasites and Insects: Aspects of Social Behavior
Disease Transmission and Networks
Climate, Cooperation and Social Evolution
The Neurobiology of Collective Behavior: Lessons from Honeybees and Ants
Foraging is fundamental for all life
Foraging: Section Overview
Application and inspiration of Optimal Foraging Theory for human endeavours outside biology
Indirect Genetic Effects
Genes, Neurones, Hormones
Cellular epigenetics and behavioral evolution
Evolution of the Nervous System in Relation to Behavior
Evolution of Behavior: Genotype to Phenotype
Evolutionary behavioral genetics
The Oxytocin System: Single Gene Effects on Social Behavior Across Species
Drosophila Behavior Genetics
Caste in Social Insects: Genetic Influences Over Caste Determination
Behavioral Genetics of Social Insects
Unicolonial Ants: Loss of Colony Identity
Behavioral genetics of dog breeds
Genetics of animal and bird migration
Major Histocompatibility Complex Genes and Mate Choice
Animal personalities and behavioural genetics
Singing Behavior in Fishes: Hormones, Neurons, and Evolution
Communication and Hormones
Sex Change in Reef Fishes: Behavior and Physiology
Male Sexual Behavior and Hormones in Non-Mammalian Vertebrates
Sexual Behavior and Hormones in Male Mammals
Female Sexual Behavior: Hormonal Basis in Non-Mammalian Vertebrates
Female Sexual Behavior and Hormones in Mammals
Hormonal and Neuromuscular Regulation of Courtship Displays
Invertebrate Hormones and Behavior
Pair Bonding, Mating Systems, and Hormones in Vertebrates
Experimental Approaches to Hormones and Behavior: Invertebrates
Aquatic Invertebrate Endocrine Disruption
Maternal Effects on Behavior
Hormones and Parental Behavior in Non-Mammalian Vertebrates
Parental Behavior and Hormones in Mammals
Tadpole Behavior and Metamorphosis
Food Intake: Behavioral Endocrinology
Aggression and Territoriality
Fight or Flight Responses
Hibernation, Daily Torpor and Estivation in Mammals and Birds: Behavioral Aspects
Circadian and Circannual Rhythms and Hormones
Wintering Strategies
Seasonality: Hormones and Behavior
Vertebrate Endocrine Disruption
Memory, Learning, Hormones and Behavior
Immune Systems and Sickness Behavior
Behavioral plasticity: the intersection of genes and environment
Hormones and Sleep
Single-cell transcriptomics and the regulation of behaviour
Social network analysis
Brain-gut axis
Field Techniques in Hormones and Behavior
Neuroethology: Methods
Sociogenomics
Sex, Genomic Imprinting and Social Evolution
Aging and Behavior in Honey Bees
Paramecium Behavioural Genetics
Endocrine and Behavioural Regulation of Water and Salt Intake in Vertebrates
Studying the Genomics of Behavior in the Genomics Era
Mating and Reproduction
Brood parasitism
Intraspecific reproductive parasitism in social insects
Reproductive Success
Propagule Behavior and Parasite Transmission
Parasites and Sexual Selection
Flexible Mate Choice
Mate Choice in Males and Females
Cryptic Female Choice
Rape, Forced and Aggressively Coerced Copulation
Compensation in Reproduction
Differential Allocation
Sex Allocation, Sex Ratios and Reproduction
Bateman׳s Principles: Original Experiment and Modern Data For and Against
Infanticide
Helping Behavior in Birds and Mammals
Sperm Competition
Gamete Sharing by “Cloacal Kissing”
Birds with Penises: Copulation Mechanics and Behavior
Nest Building in Birds
Mating Systems in Mammals
Reproductive Behavior in the Hyaenidae
Mating Systems of New World Monkeys
Mating systems of old world monkeys
Sex Changing Organisms and Reproductive Behavior
Hermaphrodite Mating Systems
Sexual Selection and Speciation
Ant, Bee and Wasp Social Evolution
Social Evolution in Termites
Social Evolution in other arthropods
Subsociality and the Evolution of Eusociality
Kin Selection and Relatedness
Division of Labor
Colony Founding in Social Insects
Queen-Queen Conflict in Eusocial Insect Colonies
Queen–Worker Conflicts Over Colony Sex Ratio
Worker Conflict and Worker Policing
Reproductive Skew, Cooperative Breeding, and Eusociality in Vertebrates: Hormones
Contact Incubation in Birds
Reproductive Behaviour and Parasites: Invertebrates
Learning, Memory and Teaching
Costs of Learning
Habitat Imprinting and Natal Habitat Preference Induction
Mate Choice and Learning
PLAY BEHAVIOR
Spatial Memory
Apes: Social Learning
Avian Social Learning
Animal Culture
Social learning in fishes
Imitation in non-human animals
Insect Social Learning
Social Learning in Non-primate mammals
Monkeys and Prosimians: Social Learning
Social Learning Theory
Vocal Learning
Social Information Use
Pavlovian learning in invertebrates
Operant learning in invertebrates
Habituation: Behavioral and Neural Mechanisms in Model Systems
Vertical Migration of Aquatic Animals
Fish Migration
Sea Turtles: Navigation and Orientation
Insect Migration
Insect Navigation
Magnetic Compasses in Insects
Amphibia Orientation and Migration
Bat Migration
Orientation Navigation and Homing in Bats
Bird Migration
Maps and Compasses
Pigeon Homing as a Model Case of Goal Oriented Navigation
Taste perception in insects
Taste Vertebrates
Invertebrate Pheromones Models for Neuroethology
Hearing Insects
Ears and hearing in vertebrates
Vibration Perception Vertebrates
Invertebrate Vision
Vision Vertebrates
Infrared Perception in Invertebrates
Electroreception in Vertebrates and Invertebrates
Active Electroreception Vertebrates
Magnetoreception - The Mechanisms, Molecules and Circuits
Neuroethology of Sound Localization in Birds
Bat Neuroethology
Vocal Acoustic Communication in Fishes Neuroethology
Acoustic Communication in Insects: Neuroethology
Leech Behavioral Choice Neuroethology
Nematode Learning and Memory Neuroethology
Neuroethology of Parasitoid Wasps
Neurobiology of pair bonding
Behavioral aspects of insect walking
Animals and us
Sentience
Assessment of welfare and needs
Indicators of good welfare
Stereotypies and Other Abnormal Behavior in Welfare Assessment
Pain in mammals; Physiology, management and assessment
Slaughter plants: behavior and welfare assessment
Stress, Health and Social Behavior
Sickness Behavior in Animals: Implications for Health and Wellness
Welfare and interactions between humans and companion animals
Race horse behavior, training and welfare
Overview of Animal Training: A Welfare Perspective
Applications of Animal Behavior to Conservation
Human impact, behavior and conservation
Conservation Behavior and Endocrinology
Noise Pollution and Conservation
Light pollution and animal conservation
Behavioral responses to climate change and chemical pollution
Wildlife in urban environments
Humans and wildlife: from conflict to coexistence
Learning and conservation
Habitat deterioration, signals and conservation
Parasitism and host behavior
Reproductive interference
The surprising link between animal behavior and the process of seed dispersal
Captive breeding, conservation and welfare
Wildlife tourism
Pig behavior and welfare
Vocal Communication Between Humans and Animals
Behaviour as a Welfare Measure in Commercial Broilers
Landmark Studies
Aplysia
Betta Splendens
Blue Tits
Boobies: Models of Evolved Family Conflict
Bowerbirds
Cockroaches: More interesting than you think
Dictyostelium, the Social Amoeba
Domestic Dogs
When expectations are thwarted: indicators and causes of frustration in hens
Grey Parrots: Studies in Avian Cognition
Honeybees
Konrad Lorenz
Locusts
Niko Tinbergen
Norway Rats
Octopus
Pheidole Ants: Sociobiology of a Highly Diverse Genus
Pigeons
Rhesus Macaques
The Sexual and Social Behavior of the Barn Swallow
Sharks and rays: Iconic species with poorly understood behaviour
Spotted Hyenas
Swordtails and Platyfishes
Threespine Stickleback
Túngara Frog: A Model for Sexual Selection and Communication
Freshwater turtles: nesting, mating and environmental sex determination
White-Crowned Sparrow
William Donald Hamilton
Wolves: families, neighborhoods and resources
Zebrafish
Karl von Frisch: Experimental Behavioral Ecologist, Ethologist and Comparative Sensory Physiologist
The evolution of intergroup conflict in animal societies
Overview Essays
Animal architecture
Survival strategies
Hormones and Behavior: Basic Concepts
Integration of Proximate and Ultimate Causes
Gene and Meme
Welfare Concepts
Neurobiology, Endocrinology and Behavior
Cognition and communication
Future of Animal Behavior: Predicting Trends
Development, Evolution and Behavior
The architecture of termite mounds’
Game theory
Methodology
Biologging and Remote-Sensing of Behavior
Data Coding, Measurement Error, and Reliability
Dominance Relationships, Dominance Hierarchies and Rankings
Endocrinology and Behavior: Methods
Ethograms, Activity Profiles, and Energy Budgets
Experimental Design: Basic Concepts
Experiment, Observation, and Modeling in the Lab and Field
Life Histories and Network Function
Sequence Analysis and Transition Models
Use of Robotics in the Study of Animal Behavior
Historical Overview
Animal Behavior: Antiquity to the Sixteenth Century
Animal Behavior: The Seventeenth to the Nineteenth Centuries
HISTORY OF THE STUDY OF ANIMAL BEHAVIOR – 20TH CENTURY
Comparative Animal Behavior - 1920 - 1973
From Ethology to behavioral biology
Behavioral Ecology and Sociobiology
Psychology of Animals
Darwin and the study of animal learning and behavior
What Animals Know About Time
Gaze and Behavior Tracking in Animal Research: Granularity, Automation, and Technological Choice
Animal Arithmetic
Animal Tool use
Categories and Concepts: Language-Related Competences in Non-Linguistic Species
Chimpanzee and bonobo
Cognitive Development in Chimpanzees
Collaborative behavior
Conflict resolution
Deception: Competition by Misleading Behavior
Emotion and Social Cognition in Primates
Empathetic Behavior
Gestural communication in the great apes
Mental Time Travel: Can Animals Recall the Past and Plan for the Future?
Metacognition and Metamemory in Non-Human Animals
Non-Elemental Learning in Invertebrates
Primate Archaeology
Comparative Thanatology: Responses to the Dying and Dead, with Special Reference to Primates
Cold-Blooded Cognition: Cognition in Testudines
Collective decision making
Avian Tool Use in the Wild
Hemispherical asymetry in animal cogntion (across species)
Acoustical Signals and Signallers – in air and water.
Chemical signaling; air, water, and on the substrate
Electric signals
Multimodal signaling
Vibrational Signals: Sounds Transmitted Through Solids
Mating signals, including advertisement and courtship
Parent-offspring Signaling
Signals in conflict resolution: conventional signals, aggression and territoriality
Individual signatures in animal groups: cetaceans
Telling Your Friends Where the Goodies are – Recruitment Signals for Food and Habitat
Signals in Insect Social Organization
Communication networks
Adjustments to Facilitate Communication in Noisy Environments
Bird vocalizations and linguistics
Rhythm and music in animal signals
Elephant communication
The Evolution of Diverse Intelligence: Comparing Corvid and Cephalopod Cognition
Elephant cognition
Behavioral Ecology
Cooperative breeding
Ecology of Fear
Isolating Mechanisms and Speciation
Methodology: Cost and Benefit Analysis
Personality and Evolution
Quantitative genetics of behavior
Specialization
An Introduction to Optimal Foraging Theory
Habitat Selection
Hunger and Satiety: Linking Mechanisms Behavior, Ecology and Evolution
Patch Exploitation
Understanding movements of foraging animals – An approach based on Optimal Foraging Theory
Food hoarding
Internal Energy Storage
How variance and risk affect foraging behaviour
Aposematism as a defence against predation
Foraging in Groups
Plant-pollinator co-evolution & Optimal Foraging Theory
Ectoparasite Behavior
Parasite-Induced Behavioral Change: Mechanisms
Avoidance of Parasites
Animal Medication: The Passive Prevention and Active Treatment of self and others
Links between Nutrition, Immunity and Infection
Reproductive Behavior and Parasites: Vertebrates
Social Behavior and Parasites
Migration in insects
Predator Evasion
Predator Avoidance: Mechanisms
Defensive Avoidance
Defensive Coloration
Group living
Anti Predatory Vigilance
Risk Allocation in Anti Predator Behavior
Defensive Chemicals
Predation risk and life histories
Defensive risk taking in Animals
Games Played by Predators and Prey
Intraguild Predation
Antipredator Benefits from Heterospecifics
Parasitoids
Crustacean Social Evolution
Microbes: Social Evolution
The Honey Bee's Dance Language
Collective Intelligence in Social Animals
Nest Site Choice in Social Insects
Group Movement
Parasites and Insects: Aspects of Social Behavior
Disease Transmission and Networks
Climate, Cooperation and Social Evolution
The Neurobiology of Collective Behavior: Lessons from Honeybees and Ants
Foraging is fundamental for all life
Foraging: Section Overview
Application and inspiration of Optimal Foraging Theory for human endeavours outside biology
Indirect Genetic Effects
Genes, Neurones, Hormones
Cellular epigenetics and behavioral evolution
Evolution of the Nervous System in Relation to Behavior
Evolution of Behavior: Genotype to Phenotype
Evolutionary behavioral genetics
The Oxytocin System: Single Gene Effects on Social Behavior Across Species
Drosophila Behavior Genetics
Caste in Social Insects: Genetic Influences Over Caste Determination
Behavioral Genetics of Social Insects
Unicolonial Ants: Loss of Colony Identity
Behavioral genetics of dog breeds
Genetics of animal and bird migration
Major Histocompatibility Complex Genes and Mate Choice
Animal personalities and behavioural genetics
Singing Behavior in Fishes: Hormones, Neurons, and Evolution
Communication and Hormones
Sex Change in Reef Fishes: Behavior and Physiology
Male Sexual Behavior and Hormones in Non-Mammalian Vertebrates
Sexual Behavior and Hormones in Male Mammals
Female Sexual Behavior: Hormonal Basis in Non-Mammalian Vertebrates
Female Sexual Behavior and Hormones in Mammals
Hormonal and Neuromuscular Regulation of Courtship Displays
Invertebrate Hormones and Behavior
Pair Bonding, Mating Systems, and Hormones in Vertebrates
Experimental Approaches to Hormones and Behavior: Invertebrates
Aquatic Invertebrate Endocrine Disruption
Maternal Effects on Behavior
Hormones and Parental Behavior in Non-Mammalian Vertebrates
Parental Behavior and Hormones in Mammals
Tadpole Behavior and Metamorphosis
Food Intake: Behavioral Endocrinology
Aggression and Territoriality
Fight or Flight Responses
Hibernation, Daily Torpor and Estivation in Mammals and Birds: Behavioral Aspects
Circadian and Circannual Rhythms and Hormones
Wintering Strategies
Seasonality: Hormones and Behavior
Vertebrate Endocrine Disruption
Memory, Learning, Hormones and Behavior
Immune Systems and Sickness Behavior
Behavioral plasticity: the intersection of genes and environment
Hormones and Sleep
Single-cell transcriptomics and the regulation of behaviour
Social network analysis
Brain-gut axis
Field Techniques in Hormones and Behavior
Neuroethology: Methods
Sociogenomics
Sex, Genomic Imprinting and Social Evolution
Aging and Behavior in Honey Bees
Paramecium Behavioural Genetics
Endocrine and Behavioural Regulation of Water and Salt Intake in Vertebrates
Studying the Genomics of Behavior in the Genomics Era
Mating and Reproduction
Brood parasitism
Intraspecific reproductive parasitism in social insects
Reproductive Success
Propagule Behavior and Parasite Transmission
Parasites and Sexual Selection
Flexible Mate Choice
Mate Choice in Males and Females
Cryptic Female Choice
Rape, Forced and Aggressively Coerced Copulation
Compensation in Reproduction
Differential Allocation
Sex Allocation, Sex Ratios and Reproduction
Bateman׳s Principles: Original Experiment and Modern Data For and Against
Infanticide
Helping Behavior in Birds and Mammals
Sperm Competition
Gamete Sharing by “Cloacal Kissing”
Birds with Penises: Copulation Mechanics and Behavior
Nest Building in Birds
Mating Systems in Mammals
Reproductive Behavior in the Hyaenidae
Mating Systems of New World Monkeys
Mating systems of old world monkeys
Sex Changing Organisms and Reproductive Behavior
Hermaphrodite Mating Systems
Sexual Selection and Speciation
Ant, Bee and Wasp Social Evolution
Social Evolution in Termites
Social Evolution in other arthropods
Subsociality and the Evolution of Eusociality
Kin Selection and Relatedness
Division of Labor
Colony Founding in Social Insects
Queen-Queen Conflict in Eusocial Insect Colonies
Queen–Worker Conflicts Over Colony Sex Ratio
Worker Conflict and Worker Policing
Reproductive Skew, Cooperative Breeding, and Eusociality in Vertebrates: Hormones
Contact Incubation in Birds
Reproductive Behaviour and Parasites: Invertebrates
Learning, Memory and Teaching
Costs of Learning
Habitat Imprinting and Natal Habitat Preference Induction
Mate Choice and Learning
PLAY BEHAVIOR
Spatial Memory
Apes: Social Learning
Avian Social Learning
Animal Culture
Social learning in fishes
Imitation in non-human animals
Insect Social Learning
Social Learning in Non-primate mammals
Monkeys and Prosimians: Social Learning
Social Learning Theory
Vocal Learning
Social Information Use
Pavlovian learning in invertebrates
Operant learning in invertebrates
Habituation: Behavioral and Neural Mechanisms in Model Systems
Vertical Migration of Aquatic Animals
Fish Migration
Sea Turtles: Navigation and Orientation
Insect Migration
Insect Navigation
Magnetic Compasses in Insects
Amphibia Orientation and Migration
Bat Migration
Orientation Navigation and Homing in Bats
Bird Migration
Maps and Compasses
Pigeon Homing as a Model Case of Goal Oriented Navigation
Taste perception in insects
Taste Vertebrates
Invertebrate Pheromones Models for Neuroethology
Hearing Insects
Ears and hearing in vertebrates
Vibration Perception Vertebrates
Invertebrate Vision
Vision Vertebrates
Infrared Perception in Invertebrates
Electroreception in Vertebrates and Invertebrates
Active Electroreception Vertebrates
Magnetoreception - The Mechanisms, Molecules and Circuits
Neuroethology of Sound Localization in Birds
Bat Neuroethology
Vocal Acoustic Communication in Fishes Neuroethology
Acoustic Communication in Insects: Neuroethology
Leech Behavioral Choice Neuroethology
Nematode Learning and Memory Neuroethology
Neuroethology of Parasitoid Wasps
Neurobiology of pair bonding
Behavioral aspects of insect walking
Animals and us
Sentience
Assessment of welfare and needs
Indicators of good welfare
Stereotypies and Other Abnormal Behavior in Welfare Assessment
Pain in mammals; Physiology, management and assessment
Slaughter plants: behavior and welfare assessment
Stress, Health and Social Behavior
Sickness Behavior in Animals: Implications for Health and Wellness
Welfare and interactions between humans and companion animals
Race horse behavior, training and welfare
Overview of Animal Training: A Welfare Perspective
Applications of Animal Behavior to Conservation
Human impact, behavior and conservation
Conservation Behavior and Endocrinology
Noise Pollution and Conservation
Light pollution and animal conservation
Behavioral responses to climate change and chemical pollution
Wildlife in urban environments
Humans and wildlife: from conflict to coexistence
Learning and conservation
Habitat deterioration, signals and conservation
Parasitism and host behavior
Reproductive interference
The surprising link between animal behavior and the process of seed dispersal
Captive breeding, conservation and welfare
Wildlife tourism
Pig behavior and welfare
Vocal Communication Between Humans and Animals
Behaviour as a Welfare Measure in Commercial Broilers
Landmark Studies
Aplysia
Betta Splendens
Blue Tits
Boobies: Models of Evolved Family Conflict
Bowerbirds
Cockroaches: More interesting than you think
Dictyostelium, the Social Amoeba
Domestic Dogs
When expectations are thwarted: indicators and causes of frustration in hens
Grey Parrots: Studies in Avian Cognition
Honeybees
Konrad Lorenz
Locusts
Niko Tinbergen
Norway Rats
Octopus
Pheidole Ants: Sociobiology of a Highly Diverse Genus
Pigeons
Rhesus Macaques
The Sexual and Social Behavior of the Barn Swallow
Sharks and rays: Iconic species with poorly understood behaviour
Spotted Hyenas
Swordtails and Platyfishes
Threespine Stickleback
Túngara Frog: A Model for Sexual Selection and Communication
Freshwater turtles: nesting, mating and environmental sex determination
White-Crowned Sparrow
William Donald Hamilton
Wolves: families, neighborhoods and resources
Zebrafish
Karl von Frisch: Experimental Behavioral Ecologist, Ethologist and Comparative Sensory Physiologist
The evolution of intergroup conflict in animal societies
Overview Essays
Animal architecture
Survival strategies
Hormones and Behavior: Basic Concepts
Integration of Proximate and Ultimate Causes
Gene and Meme
Welfare Concepts
Neurobiology, Endocrinology and Behavior
Cognition and communication
Future of Animal Behavior: Predicting Trends
Development, Evolution and Behavior
The architecture of termite mounds’
Game theory
Methodology
Biologging and Remote-Sensing of Behavior
Data Coding, Measurement Error, and Reliability
Dominance Relationships, Dominance Hierarchies and Rankings
Endocrinology and Behavior: Methods
Ethograms, Activity Profiles, and Energy Budgets
Experimental Design: Basic Concepts
Experiment, Observation, and Modeling in the Lab and Field
Life Histories and Network Function
Sequence Analysis and Transition Models
Use of Robotics in the Study of Animal Behavior
Historical Overview
Animal Behavior: Antiquity to the Sixteenth Century
Animal Behavior: The Seventeenth to the Nineteenth Centuries
HISTORY OF THE STUDY OF ANIMAL BEHAVIOR – 20TH CENTURY
Comparative Animal Behavior - 1920 - 1973
From Ethology to behavioral biology
Behavioral Ecology and Sociobiology
Psychology of Animals
Darwin and the study of animal learning and behavior
What Animals Know About Time
Gaze and Behavior Tracking in Animal Research: Granularity, Automation, and Technological Choice
- Edition: 3
- Latest edition
- Published: June 1, 2026
- Language: English
BO
Benjamin Oldroyd
Ben Oldroyd is Emeritus Professor of Behavioural Genetics at the University of Sydney. He completed a BSc in Agriculture at the University of Sydney in 1980, and a PhD on bee breeding in 1984. Ben’s research focuses on the behavioural genetics of honey bees, the evolution of social behaviour and evolution more broadly. In 2001 Ben was awarded a Doctor of Science for his contributions to the understanding of the evolution of honey bee societies. Ben is heavily involved with the Australian beekeeping industry, including helping beekeepers breed better, healthier, strains. Ben has made important contributions to our understanding of the biology of Asian honey bees and Cape bees. His book Asian Honey Bees: Biology (Harvard University Press) is the authoritative text on the subject. Ben has authored over 300 scientific papers on honey bees and stingless bees and his papers have been cited over 16,000 times.
Affiliations and expertise
Emeritus Professor of Behavioural Genetics, University of Sydney, Australia